High diversity of Endothyrella (Gastropoda, Pulmonata, Plectopylidae) in Bhutan, with a description of four new species

Recent fieldwork in Bhutan resulted in the discovery of 4 endemic species of Endothyrella that are described as new to science, viz. E. barnai Gittenberger & Sherub, E. manasensis Gittenberger & Sherub, E. pterocallus Gittenberger, Gyeltshen & Tobgay, and E. trimagnipili Gittenberger, Gyeltshen & Tobgay. This results in a total of 9 Endothyrella species for Bhutan, 7 of which are not known from elsewhere. The new species are described and additional data are presented for the Endothyrella species dealt with in an earlier paper. A revised identification key for the Bhutanese species, based on shell characters, is added. The diversity in shell size and struc­ture is discussed and 4 species groups are provisionally dis­tinguished on the basis of the structure of the genital tracts.